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A primary determinant of grain yield in barley (Hordeum vulgare L. emm. Lam) is the number of ear-bearing tillers per plant at harvest, which depends both on the production of tillers and on their subsequent survival to form ears. This three-year field study compares tiller production and survival in relation to final grain yield in three types of barley: 2-rowed winter (2rw), 6-rowed winter (6rw) and 2-rowed spring (2rs), grown in two contrasting environments. These three types differed significantly in shoot and ear number, the winter barleys showing higher tiller production, with the maximum number of tillers ranging from 798 to 2315 m−2 in 2rw, 711 to 1527 in 6rw and 605 to 1190 in 2rs. Grain yield across environments and years was strongly correlated () with the number of ears at harvest. The maximum number of shoots produced by each type of barley was inversely related to the mean temperature during the tillering phase. Tiller mortality was inversely related to the maximum shoot production, being significantly lower in barleys with less tillering capacity, i.e. the spring type (with average values of 34.3% and 42.7% in the two environments). The nment on all four characters. The entry-by-year interactions were significant for all traits, primarily because of changes in the relative genotypic differences for these traits in the four years. However, certain lines consistently ranked low or high for tiller mortality. There was a significant negative correlation between front tine tiller and grain yield in 3 out of 4 years. There was a positive correlation of highest tiller number with reproductive tiller number and with tiller mortality. Grain yield showed a nonsignificant positive correlation with maximum tiller number. The reproductive tiller number was positively correlated with grain yield. Results of this study indicate that spikeless tillers contribute negatively to grain yield and that genetic variation exists for tiller mortality in spring wheat.Timing of tiller initiation can also influence tiller mortality. Charles-Edwards (1984) concluded that self-thinning within plant communities is largely due to the lack of assimilate needed to continue growth and development within the individual stem which, in turn, can lead to a decrease in plant weight and eventually a decrease in plant yield. Some works have explored the purpose of rear tine tiller and the effects it may have on the plant as a whole and concluded tillers that abort may have benefited the plant due to assimilate and nutrient accumulation (Lupton and Pinthus, 1969; Palfi and Dezsi, 1960). However, Langer and Dougherty (1976) concluded that dead tillers had a negative effect on grain yield due to competition for assimilates and nutrients (Sharma, 1995).The second section equaling 9.12 m2 was designated for marked samples. Five plants from each plot were marked and the number of full and partial leaves on each MS and tiller were recorded along with the total number of tillers at current growth stage. This was done once a month from planting to harvest. Throughout the 2009–2010 growing season, observations were made at TRS and BC on 22 December, 28 January, 1 March, 19 March, 7 April, and 26 April. During the 2010–2011 growing season, observations were made on 7 December, 31 January, 4 March, 2 April, and 30 April. During the 2011 growing season, leaf and Garden Tiller and Cultivator counts were recorded on 9 December, 2 January, 11 February, and 3 April at TRS and 15 December, 9 January, 24 February, and 13 April at PRS. Each new and existing tiller was noted using either a black, silver, or red permanent marker to mark leaf number. Black markings represented tillers that were initiated from planting through the end of December. Silver markings represented early winter tillers that developed from the first k, silver, red) counted and hand threshed to determine the number of spikes and grain weight spike–1 for each tiller initiation period. The data for all five plants were averaged to represent values for each plot.In the destructive sample plots some samples were lost in 2011–2012. Therefore, only samples taken in 2009–2010 and 2010–2011 at TRS and BC were used in the analysis. The Proc Mixed procedure in SAS (SAS Institute, Inc.) was used to determine if there were differences in spikes m–2, kernels spike–1, weight per 100 kernels, and grain yield among site-years, mini power tiller tractor initiation period, seeding rate and N application timing. As with previous analysis, site-year, seeding rate, and N timing were treated as fixed effects; while blocks and the interactions with blocks were treated as random. When differences were detected, Fisher’s Protected LSD was used to separate means.